1. 近海和远洋环境中浮游病毒及微型生物群落生态学
2. 海洋病毒的多样性与组学信息库构建
3. 海洋病毒与赤潮生物的相互作用与关系

1. Ecological study of viral and microbial community in China Marginal Seas and in pelagic environment including Mariana Trench, Antarctic and Arctic
2. Diversity of marine viruses and omics information database construction
3. The interaction between marine viruses and red-tide